I am a PhD candidate in Biology and an MS candidate in Statistics at Louisiana State University. My research focus is the integration of mathematical biology and statistical methods in the study of plant systems. My dissertation work looks at model scale in systems ranging from the protein subnetwork to population level. 

I am also interested in STEM education. I am particularly interested in making mathematical biology more accessible to experimentalists and the general public alike.

I have a tarantula, Chinese water dragon, cat, and a few hundred Madagascar hissing cockroaches from a previous student's undergraduate lab project. Check out my tarantula digging:

Research Areas:
Computational Biology, Education and Outreach, Gene Regulation, Metabolism, Systems Biology


Dale, R., Kato, N., Wischusen, E. (Submitted, 2018) Four assessment methods to measure student gains in a graduate course on mathematical modeling in cell biology. Letters in Biomathematics. (preprint:

Dale, R., Chen, Y., He, H. (Submitted, 2018) Hierarchical modeling of the effect of pre-exposure prophylaxis on HIV in the US. (preprint:

Dale, R., Guo, B. (2018) Estimating epidemiological parameters of a stochastic differential model of HIV dynamics in the United States using hierarchical Bayesian statistics. PLoS ONE. (preprint:

Kumar, N., Dale, R., Kemboi, D., Zeringue, E. A., Kato, N., Larkin, J. C. (2018) Functional Analysis of Short Linear Motifs in the Plant Cyclin-Dependent Kinase Inhibitor SIAMESE. Plant Physiology.

Chen, Y.C., Dale, R., He, Hongyu, Le, Quoc-Anh T. Le. (2017) Posterior Estimates of Dynamic Constants in HIV Transmission Modeling. Computational and Mathematical Methods in Medicine.

Brauer, E. K., Ahsan, N., Dale, R., Kato, N., Coluccio, A. E., Piñeros, M. A., Kochian, L. V., Thelen, J. J., Popescu, S. C. (2016). The Raf-like kinase ILK1 and the high affinity K+ transporter HAK5 are required for Innate Immunity and Abiotic Stress Response. Plant Physiol. pp.00035.2016.  

Dale, R., Kato, N. (2016). Truly quantitative analysis of the firefly luciferase complementation assay. Current Plant Biology 5(2016): 57-64.

Dale, R., Ohmuro-Matsuyama, Y., Ueda, H., Kato, N. (2016). Mathematical Model of the Firefly Luciferase Complementation Assay Reveals a Non-Linear Relationship between the Detected Luminescence and the Affinity of the Protein Pair Being Analyzed. PLoS ONE 11(2): e0148256.

Kumar, N., Harashima, H., Kalve, S., Bramsiepe, J., Wang, K., Sizani, B. L., Bertrand, L. L., Johnson, M. C., Faulk, C., Dale, R., Simmons, L. A., Churchman, M. L., Sugimoto, K., Kato, N., Dasanayake, M., Beemster, G., Schnittger, A., Larkin, J. C. (2015). Functional Conservation in the SIAMESE-RELATED Family of Cyclin-Dependent Kinase Inhibitors in Land Plants. Plant Cell 27(11): 3065-3080.

Fontenot, E. B., Ditusa, S. F., Kato, N., Olivier, D. M., Dale, R., Lin, W. Y., Chiou, T. J., Macnaughtan, M. A., Smith, A. P. (2015). Increased phosphate transport of Arabidopsis thaliana Pht1;1 by site-directed mutagenesis of tyrosine 312 may be attributed to the disruption of homomeric interactions. Plant Cell Environ 38(10): 2012-2022.

Current Affiliations
Experience & Education